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5. This dataleadsto the conclusionthat eitherloss of mineralizationof the gladiusin extinct squidlike coleoidsoccurredin pre-Jurassic time, or that the gladius in pre-Jurassictime was originallymainly organicin composition. Acknowledgements: We gratefully acknowledge that the International ScienceFoundationprovided support for a field trip to the Middle Volga region in 1995, and rhe Swedish Academy of Sciencesfbr providing a grant in 2005 supportingour research. RBrBnnNcps DOGUZHAEVA, L. A. 2002.

AlSymposium- Coleoid CephalopodsThrough Time - Prague 2005 d 1 E e I a 3 ,t: Fig. 4. Schematicdrawing of the adhesiveorgan of Idiosepius sp. with its characteristiccells: co-columnarcells, grgranular cells, go-goblet cells and in-interstitial cells. Image publishedwith permissionof Norbert Cyran. Histological, histochemicaland ultrastructural methods were applied to elucidate the nature of the secretion in the epithelial cells of three ldiosepius species(1. biserialis, I. paradoxlzsand I. The adhesive organ of all speciesconsistsof four different cell types (columnar cells, granular cells, goblet cells and interstitialcells), which can be distinguished morphologically and on account of their chemistry of their secretion (Figure 4).

PEARCE,J. C. 1841. On the fossil Cephalopoda constitutingthe genusBelemnotl'tezris, PEARCE. , 2: 75-18. 47 - Coleoid CephalopodsThrough Time - Prague 2005 2"'tInternationctlS7,n7p6s1Ltm PLATE 1. EXPLANATION Pro-ostracum ultrastructure in Belemnotheutis antiquus; Oxford Clay, Jurassic; Callovian, Christian Malford, Wiltshire, England: A. SpecimenNo. SMNH 90284;B - F. No. G. Laminationin chitinous gladius(crosssection)in living squidBerryteuthismagisterBeRRv;scalebar - 30 pm. A. Middle part of the shell; the pro-ostracum(pro) is situatedbetweenthe rostrum (r) and the nacreouslayer(n) oI the conotheca;scalebar - 3Upm.

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2nd International Symposium Coleoid Cephalopods through time. hort papers. Abstract volume by Marek J., Kostak M.


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